Dominus was the channel by which the alien race named "The Arcane" conquered planet after planet. At Lucifer's command post, the Supreme One checked in with his Agent, Lucifer, and said that the time was ready for his true purpose, which was to deploy Dominus. Dominus and Lucifer were temporarily defeated by Charles Xavier, who suffered a debilitating injury in the process. Xavier's X-Men would later defeat Lucifer.
Dominus' Minions Dominus employed a number of servants (AKA the Desert Dwellers) in contending with the costumed champions who opposed its attempts to conquer the Earth. One of them was Sunstroke, a human being. The others were sentient beings created from lizards ("Gila"), cacti ("Cactus"), and rocks ("Butte") by means of the robot's alien technology. Dominus was able to create duplicates of each of the three artificially created beings.